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1967.  Heyer, W. R.  A herpetofaunal study of an ecological transect through the Cordillera de Tilarán, Costa Rica.  Copeia, 1967:259-271.

1967.  Savage, J. M., & W. R. Heyer.  Variation and distribution in the tree-frog genus Phyllomedusa in Costa Rica, Central America.  Beiträge zur Neotropischen Fauna, 5:111-131.

1968.  Heyer, W. R.  The proper name for the type-species of the genus Leptodactylus.  Copeia, 1968:160-162.

1968.  Dixon, J. R., & W. R. Heyer.  Anuran succession in a temporary pond in Colima, Mexico.  Bulletin of the Southern California Academy of Sciences, 67:129-137.

1969.  Heyer, W. R.  Studies on the genus Leptodactylus (Amphibia, Leptodactylidae).  III.  A redefinition of the genus Leptodactylus and a description of a new genus of leptodactylid frogs.  Contributions in Science, Los Angeles County Museum of Natural History, (155):1-14.

1969.  Heyer, W. R.  Studies on frogs of the genus Leptodactylus (Amphibia, Leptodactylidae).  V.  Taxonomic notes on L. latinasus, rhodonotus, romani, and wuchereri.  Herpetologica, 25:1-8.

1969.  Heyer, W. R., & P. A. Silverstone.  The larva of Leptodactylus hylaedactylus (Amphibia, Leptodactylidae).  Fieldiana:  Zoology, 51:141-145.

1969.  Heyer, W. R.  The adaptive ecology of the species groups of the frog genus Leptodactylus (Amphibia, Leptodactylidae).  Evolution, 23:421-428.

1969.  Savage, J. M., & W. R. Heyer.  The tree-frogs (Family Hylidae) of Costa Rica:  Diagnosis and distribution.  Revista de Biología Tropical, 16:1-127. 

1970.  Heyer, W. R.  Studies on frogs of the genus Leptodactylus (Amphibia, Leptodactylidae).  VI.  Biosystematics of the Melanonotus group.  Contributions in Science, Los Angeles County Museum of Natural History, (191):1-48.

1970.  Heyer, W. R.  Studies on the genus Leptodactylus (Amphibia, Leptodactylidae).  II.  Diagnosis and distribution of the Leptodactylus of Costa Rica.  Revista de Biología Tropical, 16:171-205.

1970.  Heyer, W. R., & S. Pongsapipatana.  Gliding speeds of Ptychozoon lionatum (Reptilia:  Gekkonidae) and Chrysopelea ornata (Reptilia:  Colubridae).  Herpetologica, 26:317-319.

1971.  Heyer, W. R., & J. A. Peters.  The frog genus Leptodactylus in Ecuador.  Proceedings of the Biological Society of Washington, 84:163-170.

1971.  Heyer, W. R.  Mating calls of some frogs from Thailand.  Fieldiana:  Zoology, 58:61-82.

1971.  Heyer, W. R.  Descriptions of some tadpoles from Thailand.  Fieldiana:  Zoology, 58:83-91.

1971.  Heyer, W. R.  Leptodactylus labialis.  Catalogue of American Amphibians and Reptiles, (104):1-3.

1972.  Heyer, W. R.  A new limbless skink (Reptilia:  Scincidae) from Thailand with comments on the generic status of the limbless skinks of Southeast Asia.  Fieldiana:  Zoology, 58:109-129.

1972.  Heyer, W. R.  The status of Leptodactylus pumilio Boulenger (Amphibia, Leptodactylidae) and the description of a new species of Leptodactylus from Ecuador.  Contributions in Science, Los Angeles County Museum of Natural History, (231):1-8.

1973.  Heyer, W. R., & M. S. Bellin.  Ecological notes on five sympatric Leptodactylus (Amphibia, Leptodactylidae) from Ecuador.  Herpetologica, 29:66-72.

1973.  Heyer, W. R., & K. A. Berven.  Species diversities of herpetofaunal samples from similar microhabitats at two tropical sites.  Ecology, 54:642-645.

1973.  Heyer, W. R.  Systematics of the Marmoratus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae).  Contributions in Science, Los Angeles County Museum of Natural History, (251):1-50.

1973.  Heyer, W. R.  Ecological interactions of frog larvae at a seasonal tropical location in Thailand.  Journal of Herpetology, 7:337-361.

1974.  Heyer, W. R.  Relationships of the marmoratus species group (Amphibia, Leptodactylidae) within the Subfamily Leptodactylinae.  Contributions in Science, Los Angeles County Museum of Natural History, (253):1-46.

1974.  Heyer, W. R.  Vanzolinius, a new genus proposed for Leptodactylus discodactylus (Amphibia, Leptodactylidae).  Proceedings of the Biological Society of Washington, 87:81-90. [PDF]

1974.  Heyer, W. R.  Niche measurements of frog larvae from a seasonal tropical location in Thailand.  Ecology, 55:651-656.

1974.  Heyer, W. R., & M. J. Diment.  The karyotype of Vanzolinius discodactylus and comments on usefulness of karyotypes in determining relationships in the Leptodactylus-complex (Amphibia, Leptodactylidae).  Proceedings of the Biological Society of Washington, 87:327-336. [PDF]

1975.  Heyer, W. R.  A preliminary analysis of the intergeneric relationships of the frog family Leptodactylidae.  Smithsonian Contributions to Zoology, (199):1-55. [Pt1] [Pt2]

1975.  Heyer, W. R., R. W. McDiarmid, & D. L. Weigmann.  Tadpoles, predation and pond habitats in the tropics.  Biotropica, 7:100-111.

1975.  Pyburn, W. F., & W. R. Heyer.  Identity and call of the frog, Leptodactylus stenodema.  Copeia, 1975:585-587.

1975.  Heyer, W. R.  Adenomera lutzi (Amphibia:  Leptodactylidae), a new species of frog from Guyana.  Proceedings of the Biological Society of Washington, 88:315-318. [PDF]

1976.  Straughan, I. R., & W. R. Heyer.  A functional analysis of the mating calls of the Neotropical frog genera of the Leptodactylus complex (Amphibia, Leptodactylidae).  Papéis Avulsos de Zoologia, 29:221-245.

1976.  Muedeking, M. H., & W. R. Heyer.  Descriptions of eggs and reproductive patterns of Leptodactylus pentadactylus (Amphibia:  Leptodactylidae).  Herpetologica, 32:137-139.

1976.  Heyer, W. R., & D. S. Liem.  Analysis of the intergeneric relationships of the Australian frog family Myobatrachidae.  Smithsonian Contributions to Zoology, (233):1-29. 

1976.  Heyer, W. R.  Studies in larval amphibian habitat partitioning.  Smithsonian Contributions to Zoology, (242):1-27. [PDF]

1976.  Heyer, W. R.  The presumed tadpole of Paratelmatobius lutzi (Amphibia, Leptodactylidae).  Papéis Avulsos de Zoologia, 30:133-135.

1976.  Heyer, W. R.  Notes on the frog fauna of the Amazon Basin.  Acta Amazonica, 6:369-378. [PDF]

1977.  Heyer, W. R., & M. H. Muedeking.  Notes on tadpoles as prey for naiads and turtles.  Journal of the Washington Academy of Sciences, 66:235-239. [PDF]

1977.  Heyer, W. R.  A discriminant function analysis of the frogs of the genus Adenomera (Amphibia:  Leptodactylidae).  Proceedings of the Biological Society of Washington, 89:581-592. [PDF]

1977.  Heyer, W. R., & A. S. Rand.  Foam nest construction in the leptodactylid frogs Leptodactylus pentadactylus and Physalaemus pustulosus (Amphibia, Anura, Leptodactylidae).  Journal of Herpetology, 11:225-228. [PDF]

1977.  Heyer, W. R.  Taxonomic notes on frogs from the Madeira and Purus rivers, Brasil.  Papéis Avulsos de Zoologia, 31:141-162. [PDF]

1977.  Heyer, W. R.  Variation in members of the Centrolenella eurygnatha complex (Amphibia:  Centrolenidae) from Serra do Mar and Serra da Mantiqueira, Brasil.  Papéis Avulsos de Zoologia, 32:15-33. [PDF]

1978.  Heyer, W. R.  Systematics of the fuscus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae).  Natural History Museum of Los Angeles County Science Bulletin, (29):1-85. [Pt1] [Pt2]

1979.  Heyer, W. R., & C. C. Mello.  Descriptions of the advertising calls of Cycloramphus asper and Cycloramphus dubius (Amphibia:  Leptodactylidae).  Papéis Avulsos de Zoologia, 32:193-200.

1979.  Heyer, W. R., & R. I. Crombie.  Natural history notes on Craspedoglossa stejnegeri and Thoropa petropolitana (Amphibia:  Salientia, Leptodactylidae).  Journal of the Washington Academy of Sciences, 69:17-20.

1979.  Heyer, W. R.  Systematics of the pentadactylus species group of the frog genus Leptodactylus (Amphibia:  Leptodactylidae).  Smithsonian Contributions to Zoology, (301):1-43. [PDF]

1979.  Heyer, W. R.  Annual variation in larval amphibian populations within a temperate pond.  Journal of the Washington Academy of Sciences, 69:65-74.

1980.  Heyer, W. R.  The calls and taxonomic positions of Hyla giesleri and Ololygon opalina (Amphibia:  Anura:  Hylidae).  Proceedings of the Biological Society of Washington, 93:655-661.

1981.  Maxson, L. R., W. R. Heyer, & R. D. Maxson.  Phylogenetic relationships of the Brasilian leptodactylid frog genera Craspedoglossa, Cycloramphus and Zachaenus.  Experientia, 32:144-145.

1982.  Maxson, L. R., & W. R. Heyer.  Leptodactylid frogs and the Brasilian Shield:  An old and continuing adaptive relationship.  Biotropica, 14:10-15.

1982.  Heyer, W. R., and L. R. Maxson.  Distributions, relationships, and zoogeography of lowland frogs:  The Leptodactylus complex in South America, with special reference to Amazonia.  Pages 375-388 in Ghillean T. Prance (ed.), Biological Diversification in the Tropics.  Columbia University Press. [PDF]

1982.  Heyer, W. R., and L. R. Maxson.  Neotropical frog biogeography:  Paradigms and problems.  American Zoologist, 22:397-410. [PDF]

1982.  Heyer, W. R.  Two new species of the frog genus Hylodes from Caparaó, Minas Gerais, Brasil (Amphibia:  Leptodactylidae).  Proceedings of the Biological Society of Washington, 95:377-385.

1982.  Drewry, G. E., W. R. Heyer, & A. S. Rand.  A functional analysis of the complex call of the frog Physalaemus pustulosus.  Copeia, 1982:636-645.

1982.  Heyer, W. R., C. H. Daugherty, & L. R. Maxson.  Systematic resolution of the genera of the Crinia complex (Amphibia:  Anura:  Myobatrachidae).  Proceedings of the Biological Society of Washington, 95:423-427.

1983.  Wassersug, R. J., & W. R. Heyer.  Morphological correlates of subaerial existence in leptodactylid tadpoles associated with flowing water.  Canadian Journal of Zoology, 61:761-769.

1983.  Heyer, W. R.  Clarification of the names Rana mystacea Spix, 1824, Leptodactylus amazonicus Heyer, 1978 and a description of a new species, Leptodactylus spixi (Amphibia:  Leptodactylidae).  Proceedings of the Biological Society of Washington, 96:270-272. [PDF]

1983.  Heyer, W. R. Variation and systematics of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae).  Arquivos de Zoologia, 30:235-339.

1983.  Heyer, W. R., & L. R. Maxson.  Relationships, zoogeography, and speciation mechanisms of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae).  Arquivos de Zoologia, 30:341-373. [PDF]

1983.  Heyer, W. R.  Notes on the frog genus Cycloramphus (Amphibia:  Leptodactylidae), with descriptions of two new species.  Proceedings of the Biological Society of Washington, 96:548-559.

1983.  Heyer, W. R., & W. F. Pyburn.  Leptodactylus riveroi, a new frog species from Amazonia, South America (Anura:  Leptodactylidae).  Proceedings of the Biological Society of Washington, 96:560-566. [PDF]

1983.  Crombie, R. I., & W. R. Heyer.  Leptodactylus longirostris (Anura:  Leptodactylidae):  Advertisement call, tadpole, ecological and distributional notes.  Revista Brasileira de Biologia, 43:291-296. [PDF]

1984.  Heyer, W. R.  Variation, systematics, and zoogeography of Eleutherodactylus guentheri and closely related species (Amphibia:  Anura:  Leptodactylidae).  Smithsonian Contributions to Zoology, (402):1-42. [PDF]

1984.  Heyer, W. R.  The systematic status of Adenomera griseigularis Henle, with comments on systematic problems in the genus Adenomera (Amphibia:  Leptodactylidae).  Amphibia-Reptilia, 5:97-100. [PDF]

1985.  Heyer, W. R.  Taxonomic and natural history notes on frogs of the genus Centrolenella (Amphibia:  Centrolenidae) from southeastern Brasil and adjacent Argentina.  Papéis Avulsos de Zoologia, 36:1-21.

1985.  Heyer, W. R.  New species of frogs from Boracéia, São Paulo, Brazil.  Proceedings of the Biological Society of Washington, 98:657-671.

1985.  Vanzolini, P. E., & W. R. Heyer.  The American herpetofauna and the interchange.  Pages 475-487 in F. G. Stehli & D. S. Webb (editors), The Great American Biotic Interchange.  Plenum Press. [PDF]

1986.  Heyer, W. R., & R. B. Cocroft.  Descriptions of two new species of Hylodes from the Atlantic Forests of Brazil (Amphibia:  Leptodactylidae).  Proceedings of the Biological Society of Washington, 99:100-109.

1988.  Maxson, L. R., & W. R. Heyer.  Molecular systematics of the frog genus Leptodactylus (Amphibia:  Leptodactylidae).  Fieldiana:  Zoology, New Series, (41):1-13. [PDF]

1988.  Heyer, W. R.  A notable collection of Cycloramphus (Amphibia:  Leptodactylidae) from Bahia, Brasil, with a description of a new species (Cycloramphus migueli).  Proceedings of the Biological Society of Washington, 101:151-154.

1988.  Cocroft, R. B., & W. R. Heyer.  Notes on the frog genus Thoropa (Amphibia:  Leptodactylidae) with a description of a new species (Thoropa saxatilis).  Proceedings of the Biological Society of Washington, 101:209-220.

1988.  Wassersug, R. J., & W. R. Heyer.  A survey of internal oral features of leptodactyloid larvae (Amphibia:  Anura).  Smithsonian Contributions to Zoology, (457):1-99. [Pt1] [Pt2] [Pt3] [Pt4]

1988.  Heyer, W. R., A. S. Rand, C. A. G. Cruz, & O. L. Peixoto.  Decimations, extinctions, and colonizations of frog populations in southeast Brazil and their evolutionary implications.  Biotropica, 20:230-235.

1988.  Heyer, W. R., & P. E. Vanzolini.  Introduction and summary of a workshop on Neotropical distribution patterns held 12-16 January 1987.  Pages 7-18 in P. E. Vanzolini & W. R. Heyer (editors), Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987.  Academia Brasileira de Ciências.

1988.  Heyer, W. R.  On frog distribution patterns East of the Andes.  Pages 245-273 in P. E. Vanzolini & W. R. Heyer (editors), Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987.  Academia Brasileira de Ciências. [PDF]

1989.  Heyer, W. R., & A. J. Wolf.  Phylalaemus crombiei (Amphibia:  Leptodactylidae), a new frog species from Espírito Santo, Brazil with comments on the P. signifer group.  Proceedings of the Biological Society of Washington, 102:500-506.

1990.  Heyer. W. R., A. S. Rand, C. A. G. Cruz, O. L. Peixoto, & C. E. Nelson.  Frogs of Boracéia.  Arquivos de Zoologia, 31:231-410. 

1991.  Heyer, W. R., & L. M. Hardy.  A new species of frog of the Eleutherodactylus lacrimosus assembly from Amazonia, South America (Amphibia:  Anura:  Leptodactylidae).  Proceedings of the Biological Society of Washington, 104:436-447. 

1992.  Dubois, A. & W. R. Heyer.  Leptodactylus labialis, the valid name for the American White-Lipped Frog (Amphibia:  Leptodactylidae).  Copeia, 1992:584-585. [PDF]

1993.  García Pérez, L., & W. R. Heyer.  Description of the advertisement call and resolution of the systematic status of Leptodactylus gracilis delattini Müller, 1968 (Amphibia:  Leptodactylidae).  Proceedings of the Biological Society of Washington, 106:51-56. [PDF]

1994.  Heyer, W. R.  Variation within the Leptodactylus podicipinus-wagneri complex of frogs (Amphibia:  Leptodactylidae).  Smithsonian Contributions to Zoology, (546):1-124. [PDF]

1994.  Heyer, W. R.  Thread bobbins.  Pages 153-155 in W. R. Heyer, M. A. Donnelly, R. W. McDiarmid, L.-A. C. Hayek, & M. S. Foster (editors), Measuring and Monitoring Biological Diversity:  Standard Methods for Amphibians.  Smithsonian Institution Press.

1994.  Fellers, G. M., C. A. Drost, & W. R. Heyer.  Handling live amphibians.  Pages 275-276 in W. R. Heyer, M. A. Donnelly, R. W. McDiarmid, L.-A. C. Hayek, & M. S. Foster (editors), Measuring and Monitoring Biological Diversity:  Standard Methods for Amphibians.  Smithsonian Insititution Press. [PDF]

1994.  Heyer, W. R.  Recording frog calls.  Pages 285-287 in W. R. Heyer, M. A. Donnelly, R. W. McDiarmid, L.-A. C. Hayek, & M. S. Foster (editors), Measuring and Monitoring Biological Diversity:  Standard Methods for Amphibians.  Smithsonian Institution Press. [PDF]

1994.  Jacobs, J. F., & W. R. Heyer.  Collecting tissue for biochemical analysis.  Pages 299-301 in W. R. Heyer, M. A. Donnelly, R. W. McDiarmid, L.-A. C. Hayek, & M. S. Foster (editors), Measuring and Monitoring Biological Diversity:  Standard Methods for Amphibians.  Smithsonian Institution Press. [PDF]

1994.  Heyer, W. R.  Hyla benitezi (Amphibia:  Anura:  Hylidae):  First record for Brazil and its biogeographical significance.  Journal of Herpetology, 28:497-499.

1995.  Heyer, W. R., & V. R. Morales.  The advertisement call of the leptodactylid frog Leptodactylus griseigularis.  Amphibia-Reptilia, 16:91-92. [PDF]

1995.  Heyer, W. R., & C. Gascon.  Collection notes and call characteristics for Ischnocnema quixensis and Phyzelaphryne miriamae (Amphibia:  Anura:  Leptodactylidae).  Journal of Herpetology, 29:304-307.

1995.  Cardoso, A. J., & W. R. Heyer.  Advertisement, aggressive, and possible seismic signals of the frog Leptodactylus syphax (Amphibia, Leptodactylidae).  Alytes, 13:67-76. [PDF]

1995.  Heyer, W. R.  South American rocky habitat Leptodactylus (Amphibia:  Anura:  Leptodactylidae) with description of two new species.  Proceedings of the Biological Society of Washington, 108:695-716. [PDF]

1996.  Heyer, W. R., J. M. García-López, & A. J. Cardoso.  Advertisement call variation in the Leptodactylus mystaceus species complex (Amphibia:  Leptodactylidae) with a description of a new sibling species.  Amphibia-Reptilia, 17:7-31. [PDF]

1996.  Heyer, W. R., R. de Sá, J. R. McCranie, & L. D. Wilson.  Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae):  Natural history notes, advertisement call, and relationships.  Herpetological Natural History, 4:169-174. [PDF]

1997.  Savage, J. M., & W. R. Heyer.  Digital webbing formulae for anurans:  A refinement.  Herpetological Review, 28:131.

1997.  Heyer, W. R.  Geographic variation in the frog genus Vanzolinius (Anura: Leptodactylidae).  Proceedings of the Biological Society of Washington, 110:338-365. [PDF]

1998.  Heyer, W. R.  The relationships of Leptodactylus diedrus (Anura, Leptodactylidae).  Alytes, 16:1-24. [PDF]

1998.  Kress, W. J., W. R. Heyer, P. Acevedo, J. Coddington, D. Cole, T. L. Erwin, B. J. Meggers, M. Pogue, R. W. Thorington, R. P. Vari, M. J. Weitzman, & S. H. Weitzman.  Amazonian biodiversity:  Assessing conservation priorities with taxonomic data.  Biodiversity and Conservation, 7:1577-1587. [PDF]

1999.  Heyer, W. R., & A. M. Muñoz.  Validation of Eleutherodactylus crepitans Bokermann, 1965, notes on the types and type locality of Telatrema [sic] heterodactylum Miranda-Ribeiro, 1937, and description of a new species of Eleutherodactylus from Mato Grosso, Brazil (Amphibia: Anura: Leptodactylidae).  Proceedings of the Biological Society of Washington, 112:1-18.

1999.  Heyer, W. R.  A new genus and species of frog from Bahia, Brazil (Amphibia: Anura: Leptodactylidae) with comments on the zoogeography of the Brazilian campos rupestres.  Proceedings of the Biological Society of Washington, 112:19-39.

2000.  Heyer, W. R.  The lectotype of Cystignathus hylodes Reinhardt and Lütken, 1862.  Journal of Herpetology, 34:150-153. [PDF]

2000.  Heyer, W. R., & C. M. Carvalho.  Calls and calling behavior of the frog Leptodactylus natalensis (Amphibia: Anura: Leptodacytlidae).  Proceedings of the Biological Society of Washington, 113:284-290. [PDF]

2000.  (“1999”).  Heyer, W. R., J. Coddington, W. J. Kress, P. Acevedo, D. Cole, T. L. Erwin, B. J. Meggers, M. G. Pogue, R. W. Thorington, R. P. Vari, M. J. Weitzman, & S. H. Weitzman.  Amazonian biotic data and conservation decisions.  Ciência e Cultura, Journal of the Brazilian Association for the Advancement of Science, 51:372-385. [PDF]

2000.  Heyer, W. R., & C. M. Carvalho.  The enigmatic advertisment call of Eleutherodactylus ramagii (Amphibia: Anura: Leptodactylidae).  Amphibia-Reptilia, 21:117-121.

2000. Heyer, W. R., & A. S. Thompson.   Leptodactylus rugosus.  Catalogue of American Amphibians and Reptiles, (708):1-5. [PDF]

2001.  Hayek, L.-A. C., W. R. Heyer, & C. Gascon.  Frog morphometrics:  A cautionary tale.  Alytes, 18:153-177. [PDF]

2001.  Heyer, W. R., & M. M. Heyer.  Leptodactylus lithonaetes.   Catalogue of American Amphibians and Reptiles, (723):1-3. [PDF]

2001.  Wynn, A., & W. R. Heyer.  2001.  Do geographically widespread species of tropical amphibians exist?  An estimate of genetic relatedness within the Neotropical frog Leptodactylus fuscus (Schneider 1799) (Anura Leptodactylidae).  Tropical Zoology, 14:255-285. [PDF]

2001.  Carey, C., W. R. Heyer, J. Wilkinson, R. A. Alford, J. W. Arntzen, T. Halliday, L. Hungerford, K. R. Lips, E. M. Middleton, S. A. Orchard, & A. S. Rand.  Amphibian declines and environmental change:  Use of remote-sensing data to identify environmental correlates.  Conservation Biology, 15:903-913.

2002.  Heyer, W. R.  Leptodactylus fragilis, the valid name for the Middle American and northern South American white-lipped frog (Amphibia: Leptodactylidae).  Proceedings of the Biological Society of Washington, 115:321-322. [PDF]

2002.  Heyer, W. R., & M. M. Heyer.  Leptodactylus elenae.  Catalogue of American Amphibians and Reptiles, (742):1-5. [PDF]

2002.  Heyer, W. R., R. O. de Sá, & S. Muller.  Leptodactylus silvanimbus.  Catalogue of American Amphibians and Reptiles, (743):1-3. [PDF]

2003.  Heyer, W. R., & Y. R. Reid.  Does advertisement call variation coincide with genetic variation in the genetically diverse frog taxon currently known as Leptodactylus fuscus (Amphibia: Leptodactylidae)?  Anais da Academia Brasileira de Ciências, 75:39-54. [PDF]

2003.  Heyer, M. M., W. R. Heyer, S. Spear, & R. O. de Sá.  Leptodactylus mystacinus.  Catalogue of American Amphibians and Reptiles, (767):1-11. [PDF]

2003.  Heyer, W. R., & F. A. Juncá.  Leptodactylus caatingae, a new species of frog from eastern Brazil (Amphibia: Anura: Leptodactylidae).  Proceedings of the Biological Society of Washington, 116(2):317-329. [PDF]

2004.  Heyer, W. R., & M. M. Heyer.  Leptodactylus furnarius.  Catalogue of American Amphibians and Reptiles, (785):1-5. [PDF]

2005.  Hayek, L. C., & W. R. Heyer.  Determining sexual dimorphism in frog measurement data:  Integration of statistical significance, measurement error, effect size and biological significance.  Anais da Academia Brasileira de Ciências, 77(1):45-76. [PDF]

2005.  Heyer, W. R., R. O. de Sá, & S. Muller.  On the enigmatic distribution of the Honduran endemic Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae).  Pages 81-101 in M. A. Donnelly, B. I. Crother, C. Guyer,  M. H. Wake, & M. E. White (editors), Ecology and Evolution in the Tropics:  A Herpetological Perspective.  The University of Chicago Press: Chicago & London.  xv + 675 pages. [PDF]

2005.  Heyer, W. R., & J. B. Murphy.  Declining Amphibian Populations Task Force.  Pages 17-21in  M. J. Lannoo (editor), Amphibian Declines.  The Conservation Status of United States Species.  University of California Press:  Berkeley & Los Angeles.  xii + 1094 pages. [PDF]

2005.  Heyer, W. R.  Leptodactylus fragilis Brocchi, 1877.  White-lipped frog.  Pages 500-501 in M. J. Lannoo (editor), Amphibian Declines.  The Conservation Status of United States Species.  University of California Press:  Berkeley & Los Angeles.  xxi + 1094 pages. [PDF]

2005.  Caramaschi, U., R. O. de Sá, & W. R. Heyer.  Common names for the frog genus Leptodactylus (Amphibia, Anura, Leptodactylidae).  Herpetological Review, 36(2):119-120. [PDF]

2005.  Heyer, W. R., & R. I. Crombie.  Leptodactylus lauramiriamae, a distinctive new species of frog (Amphibia: Anura: Leptodactylidae) from Rondônia, Brazil.  Proceedings of the Biological Society of Washington, 118(3):590-595. [PDF]

2005.  Heyer, W. R., R. O. de Sá, & A. Rettig.  Sibling species, advertisement calls, and reproductive isolation in frogs of the Leptodactylus pentadactylus species cluster (Amphibia, Leptodactylidae).  In N. Ananjeva & O. Tsinenko (editors), Herpetologica Petropolitana.  Proceedings of the 12th Ordinary General Meeting of the Societas Europaea Herpetologica, August 12-16, 2003, St. Petersburg, Russia.  Russian Journal of Herpetology, 12(Supplement):35-39. [PDF]

2005.  de Sá, R. O., W. R. Heyer, & A. Camargo.  Are Leptodactylus didymus and L. mystaceus phylogenetically sibling species (Amphibia, Anura, Leptodactylidae)?  ).  In N. Ananjeva & O. Tsinenko (editors), Herpetologica Petropolitana.  Proceedings of the 12th Ordinary General Meeting of the Societas Europaea Herpetologica, August 12-16, 2003, St. Petersburg, Russia.  Russian Journal of Herpetology, 12(Supplement):90-92. [PDF]

2005.  Heyer, W. R.  Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern  South America, and Amazonia.  Arquivos de Zoologia, 37(3):269-348. [PDF]

2005.  Brandão, R. A., and W. R. Heyer.  The complex calls of Leptodactylus pustulatus (Amphibia, Anura, Leptodactylidae).  Amphibia-Reptilia, 26(4):566-570.[PDF]

2006.  Camargo, A., R. O. de Sá, and W. R. Heyer.  Phylogenetic analyses of mtDNA sequences reveal three cryptic lineages in the widespread neotropical frog Leptodactylus fuscus (Schneider, 1799) (Anura, Leptodactylidae).  Biological Journal of the Linnean Society, 87:325-341. [PDF]

2006.  Heyer, W. R., & N. J. Scott, Jr.  The advertisement call of Leptodactylus laticeps (Amphibia, Anura, Leptodactylidae):  Predatory aural luring?  Herpetological Natural History, 9(2):189-194. [PDF]

2006.  Heyer, W. R., & M. M. Heyer.  Leptodactylus knudseni.  Catalogue of American Amphibians and Reptiles, (807):1-12. [PDF]

2006.  Heyer, W. R., & M. M. Heyer.  Leptodactylus natalensis.  Catalogue of American Amphibians and Reptiles, (808):1-5. [PDF]

2006 (“2005”).  de Sá, R. O., W. R. Heyer, & A. Camargo.  A phylogenetic analysis of Vanzolinius Heyer, 1974 (Amphibia, Anura, Leptodactylidae):  Taxonomic and life history implications.  Arquivos do Museu Nacional, Rio de Janeiro, 63(4):707-726. [PDF]

2006.  Heyer, M. M., W. R. Heyer, & R. O. de Sá.  Leptodactylus fragilis.  Catalogue of American Amphibians and Reptiles, (830):1-26. [PDF]

2006.  Yanek, K., W. R. Heyer, & R. O. de Sá.  Genetic resolution of the enigmatic Lesser Antillean distribution of the frog Leptodactylus validus (Anura, Leptodactylidae).  South American Journal of Herpetology, 1(3):192-201. [PDF]

2007.  Ponssa, M. L., and W. R. Heyer.  Osteological characterization of four putative species of the genus Adenomera (Anura: Leptodactylidae), with comments on intra- and interspecific variation.  Zootaxa, (1403):37-54. [PDF]

 

 

Non-Peer Reviewed Publications


1970.  Heyer, W. R.  Report No. 12:  Field work for a study of amphibians and reptiles of the ASRCT Sakaerat Experiment Station, Amphoe Pak Thong Chai, Changwat Nakhon Ratchasima.  Cooperative Research Programme No. 27, Tropical Environmental Data (TREND) Ecosystem Study of Tropical Dry-Evergreen Forest, Research Project 27/1, Description of Tropical Dry-Evergreen Forest Ecosystem.  25 pages.

1971.  Heyer, W. R.  The tadpole of Rana pileata.  Herpetological Review, 3:95.

1973.  Heyer, W. R.  [Review of] Evolutionary relationships, osteology, and zoogeography of leptodactyloid frogs by John D. Lynch.  HISS Titles and Reviews, 1:35-36.

1976.  Heyer, W. R.  [Review of] Evolution of the Desert Biota, edited by David W. Goodall.  Copeia, 1976:614-615.

1978.  Heyer, W. R.  [Review of] A Biogeographical Analysis of the Chihuahuan Desert through its Herpetofauna, by David J. Morafka.  Herpetological Review, 9:50-51.

1984.  Heyer, W. R.  [Review of] Advances in Herpetology and Evolutionary Biology:  Essays in Honor of Ernest E. Williams, edited by A. G. J. Rhodin & K. Miyata.  Science, 223:162.

1995.  Bishop, C., D. Bradford, G. Casper, S. Corn, S. Droege, G. Fellers, P. Geissler, D. M. Green, R. Heyer, M. Lannoo, D. Larson, D. Johnson, R. McDiarmid, J. Sauer, B. Shaffer, H. Whiteman, & H. Wilbur.  A proposed North American amphibian monitoring program.  Canadian Association of Herpetologists, Bulletin, 9:6-13.

1995.  Bertoluci, J.; & W. R. Heyer.  Boracéia update.  Froglog, (14):3.

1996.  Heyer, R.  Vanishing amphibians:  A real problem?  World Conservation, 1/96:10-11.

1996.  Heyer, R.  Declining Amphibian Populations Task Force.  Species, (26-27):149.

1997.  Halliday, T. R., & W. R. Heyer.  The case of the vanishing frogs.  Technology Review, 100:56-63.

1997.  Heyer, R.  Declining Amphibian Populations Task Force.  Species, (29):66.

1999.  Heyer, R., & T. Halliday.  Declining Amphibian Populations Task Force.  Past, present, and future.  Species, (31-32):117-118.

2000.  Halliday, T., R. Heyer, & J. Hanken.  Declining Amphibian Populations Task Force.  Report on activities over the past triennium and plans for the future.  Species, (34):145-146.

2000.  Heyer, W. R.  Amphibian declines and extinctions:  The situation in Latin America.  Pages 47-55 in J. E. Péfaur (editor), Ecología Latinoamericana.  Actas del III Congreso Latinoamericano de Ecología.  Consejo de Desarrollo Científico, Humanístico y Tecnológico & Consejo de Publicaciones de la Universidad de Los Andes: Mérida. 

2002.  Heyer, W. R., & U. Caramaschi.  Comment on the proposed conservation of the specific name of Leptodactylus chaquensis Cei, 1950 (Amphibia, Anura) (Case 3172; see BZN 58:116-118).  Bulletin of Zoological Nomenclature, 59:44-45.

2003.  Hayek, L.-A. C., & R. Heyer.  Adaptive cluster sampling for amphibians.  Froglog, (55):2-3.

2004.  Heyer, W. R.  Historical Perspectives:  Paulo Emílio Vanzolini.  Copeia, 2004:184-189. [PDF]

2006.  Heyer, W. R.  Obituary.  In celebration of A. Stanley Rand.  Phyllomedusa 5(1):8-10. [PDF]

2006.  Heyer, W. R., U. Caramaschi, & R. O. de Sá.  Rana ocellata Linnaeus, 1758 (currently Leptodactylus ocellatus; Amphibia, Anura:  Proposed conservation of usage of the specific name by the designation of a neotype.  Bulletin of Zoological Nomenclature, 63(3):184-186. [PDF]

 

Edited Volumes


1988.  Vanzolini, P. E., & W. R. Heyer (editors).  Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987.  Academia Brasileira de Ciências.  488 pages.

1994.  Heyer, W. R., M. A. Donnelly, R. W. McDiarmid, L.-A. C. Hayek, & M. S. Foster (editors), Measuring and Monitoring Biological Diversity:  Standard Methods for Amphibians.  Smithsonian Institution Press.  364 pages + xix.

 

 

 

 

 

 

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